Gramma, the beloved Galápagos tortoise and oldest resident of the San Diego Zoo, has passed away after more than a century. Arriving in the 1930s, she became a foundational figure in the zoos conservation efforts and a quiet celebrity known for her sweet personality. Her longevity and calm presence offered a unique connection to a bygone era, and her legacy as a gentle giant will continue to inspire awe and appreciation for these magnificent creatures.
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